The Development Of DorothyThe Movie The Wizard Of Oz Opens On A Farm I

The Development of DorothyThe film The Wizard of Oz opens on a homestead in Kansas. The lead character is a twelve-year-old young lady named Dorothy. Dorothy exists in a world restricted by matured convictions and fears, which make up an extraordinary piece of her existence. In Kansas, Dorothy is a twelve-year-old young lady, with twelve-year-old needs and feelings. I feel that the film is a gadget by which we can break down Dorothy in all perspectives, her cognizant, pre-cognizant, and oblivious. I accept that Sigmund Freud's standards on the structure of character and dreaming will help in understanding Dorothy's development, needs, and needs. Dorothy in her cognizant state battles to be heard and comprehended. She is troubled and unfit to confer her sentiments of upset and trepidation to an open ear. Her id shows itself with the longing for guaranteed satisfaction, animated by her dread that the character Ms. Gorge will obliterate Toto. Similarly as Dorothy can account for herself to her sustaining Aunt and Uncle, the Gulch character enters. Gorge takes Toto from Dorothy, with no regret, stuffs the canine in a bushel, and leaves. Dorothy rushes to her space to sob and twists up into a fetal position. The fetal position is her oblivious want for calming and having a place. Her sobbing is stopped by the arrival of her canine Toto, who had the option to outsmart Gulch. Promptly Dorothy starts preparing thoughts of fleeing in order to shield her dearest Toto from risk. Still persuaded by id like instinct, Dorothy escapes with Toto. I accept that her id is solid, but at the same time is being checked by conscience. Fleeing is id like, however securing Toto is self image's capacity to keep id in check.A meandering educator intrudes on Dorothy's mass migration from her ranch. Educator Marvel produces sentiments of miracle and solace in Dorothy. She trusts he will have the option to spare her and Toto from the underhanded grasps of Ms. Gorge. Wonder sits Dorothy dow n and endeavors to peruse her fortune in his gem ball. He brings Dorothy to the acknowledgment that fleeing was not right, and in doing so she hurt her Auntie. I accept that since she is so youthful, Dorothy needs an outer superego to give her a proportionate measure of blame for what she is doing. She understands that she wasn't right and runs home.Dorothy's arrival home is trailed by a tempest conveying a tornado. The tornado may connote a leftover strife in Dorothy's oblivious psyche, yet dually goes about as a transport for Dorothy. She comes back to her home to discover everybody has vanished into the tempest basement. Scared by the tempest, she comes back to her room and falls down on her bed. Dorothy is thumped oblivious by the window shade as it flies open and hits her in the head. She is propelled into a psychological oblivious state. The tornado turns numerous natural appearances and things past her window until at long last she observes Ms. Ravine's change from old maid, to the fiendish witch of the west. Out of nowhere everything is calm and back to typical, or so it appears. Dorothy cautiously strolls to the front entryway and opens it to discover the heaven that is Oz. The entryway opening connotes her total progress from cognizant to oblivious. The twister was the power that stirred her obviousness and inundated her in lala land - Oz. Dorothy rises up out of her home to be told, by the great witch of the north Glinda, that the evil witch of the east has been crushed by the Kansas residence. We are continually making and changing our existence, as a dreamscape and I accept that the demise of the abhorrent witch is Dorothy's method of adapting to her past dread of Gulch. The imaginative potential inside the fantasy effectively changes clashing material and uncertain emotions, into encouraging or justifiable images.Dorothy is currently gone up against with a craving to get back. Before she can loosen up she is by and by acquainted with one more ima ge of malice, the Wicked Witch of the west. After dangers from the insidious witch, Dorothy is prompted by the great witch to request the assistance of the Wizard of Oz. She is advised by local people to

Theories of Public Policy and Administration Coursework

Hypotheses of Public Policy and Administration - Coursework Example The establishing and supporting scholars go further to guarantee that liberator change in social orders is significant bearing the way that contemporary social presence neglects to address the fundamental issues required by the enlightened residents. In a transition to disentangle this hypothesis, the establishing scholars of the Critical Social hypothesis hold that life in the advanced world is similarly nonsensical as in the postmodern social orders. Indeed, even with the extensive and expanding majority rules systems in the advanced social orders, mindlessness keeps on persevering as people and countries obliterate one another. Social judgments, misery, bondage, debasement that makes a few networks proceed in destitution as different succeeds characterize just however a couple of attributes of the cutting edge world that are under analysis by the establishing scholars. As saw By Leonardo (2004), the Critical social hypothesis makes social investigation by concentrating on the veri fiable occasions and their longitudinal changes reserved and noted before the finish of work or change. Another hypothesis of Public approach and organization is the Complexity hypothesis created and researched by George Cowan of the Santa Fe Institute in I984 and Stephen Wolfram of the Center for Complexity in Illinois in 1986 (Klijn, 2007). As per the proposers of the hypothesis, the entire or whole framework is effortlessly overseen and exceptionally productive contrasted with the entirety of the parts or work done by singular pieces of the framework. As the portrayal of the hypothesis, complex frameworks incorporate various collaborating leaves behind each carrying on in the neighborhood setting as per the laws, powers, and decides that abrogate the framework. In straightforward terms, the intricate hypothesis holds that frameworks are self-sorting out and produce results that when followed can't prompt the commitment of each part in guaranteeing the resultant result.

Essay Writing Topics For High School

Essay Writing Topics For High SchoolThe essay writing topics for high school are rather simple and quite easy to pick. These topics can actually be classified into two separate categories, the academic topics and the writing projects that a student wants to do. So you must have all these things right before you start your project.For academics, there are no specific high school essay topics for high school. There are a number of subjects that your assignment can contain which is already specified by the subject matter of your high school curriculum. These subjects may be; Language Arts, Social Studies, Science and Math.When it comes to writing project, you can also choose among these subjects but the other subjects are really not included in this category. This is mainly because a writing project is usually an outline, a brainstorming session to plan out your project. This can be done with any type of topic, however, if you're going to write an essay on history or religion, it is mor e than likely that your subject should have an easier introduction to it.These high school topics for essays are basically targeted towards a specific audience that is associated with the main topic of the essay. For example, if you're going to write an essay on reading, the kind of audience you should write for will be younger students or those who are already in the college entrance examinations. These are the groups that need an easy and simple essay with all the important information in them so that they can be able to understand their learning experiences well enough.On the other hand, if you want to write an essay on math, then there are many essay topics for high school that you can use, such as, History of Mathematics. This topic can be used for a student who has good knowledge of math, especially when the age of the student is in the higher grades. In addition, there are a lot of subjects like; Science, Engineering, Biology, Geography, Technology, Arts and many others.If yo u are new to writing an essay, then you should first get in touch with a professional to help you out. These essay writing topics for high school can be a very helpful reference and guide that can help you out in your writing process and can be considered as a partner in your learning process.You can even consider hiring a tutor who can assist you in formulating and choosing the appropriate essay topics for high school. You can also contact your school counselor or your English teacher to help you out with selecting the right essay topics for high school.So when you are choosing essay writing topics for high school, all you have to do is to take note of what your knowledge and interests are. Remember that no matter what topic you choose, the only way you will be able to successfully finish your assignment is by studying well and by having enough patience.

Cleveland Bay Horse - Free Essay Example

Sample details Pages: 30 Words: 9101 Downloads: 1 Date added: 2017/06/26 Category Statistics Essay Did you like this example? Chapter 3: Demographics, Genetic Diversity and Breed Structure of the Cleveland Bay Horse Assessed By Pedigree Analysis 1. Introduction In the previous chapters we have been introduced to the importance of understanding the nature of livestock biodiversity in order to identify and develop strategies to counter its erosion. In recent years much effort has been put into understanding breed diversity and structure through molecular methods. Whilst there can be no doubt this has advanced knowledge, it is only very recently that the trend in research has come full circle back to appreciation of the importance of analysis of pedigree data, to uncover the nature of founder contributions and the levels of inbreeding brought about through historic breeding practices. With the increase in availability of appropriate software there has been a growth in the number of livestock breeds being successfully described in genetic terms, including specific closed populations of different equine breeds such as the Lipizzan(Zechner et al., 2002), Polish Arabian (Glazewska and Jezierski, 2004), Andalusian(Valera et al., 2005) and Fresian(Ducro et al., 2006). Don’t waste time! Our writers will create an original "Cleveland Bay Horse" essay for you Create order In this chapter we look in detail at what can be learned from analysis of the Cleveland Bay Horse Society Studbook Data, and gain a more complete understanding of why the breed is placed on the Critically Endangered watch-list of the Rare Breeds Survival Trust. 2. Materials and Methods 2.1. Data Acquisition The Cleveland Bay Horse Society published its first studbook in 1884. Volume One contained retrospective pedigrees of 567 stallions foaled prior to January 1880. The Society has maintained its studbook since that time, now comprising 38 volumes and containing pedigree information on 5757 animals. Since Volume III, which included retrospective data for numerous mares as well as additional stallions, the studbook has remained effectively closed, and now requires 8 generation pedigrees for animals to be eligible for inclusion in the pure bred register. In addition to the pure register, since 1934, the Society has also maintained a Grading Register, in which mares of Cleveland Type can be listed and be used in an upgrading scheme. This scheme was originally conceived at a time when breed numbers were heavily depleted as a result of losses in the Great War of 1914 1918, and breeders recognised that there were still unregistered mares of probable Cleveland Bay origin of farms in the North East of England. In recent years very few mares have been added to the Grading Register, numbering single s in the last decade. In 1985 the Society published its 29th volume the Centenary studbook, which contained a summary of all animals recorded in previous volumes. Until the mid 1990s all society studbook records were in paper format, with no electronic data available. The data for the present study was extracted from Volume 29 and the subsequent 9 editions of the studbook. The information was digitised in Filemaker database format, listing the unique identities of each animal, along with those of the two parent animals (where known); the sex of each animal and date of birth. Subsequent to this first complete digitisation, the electronic data was made available to the Cleveland Bay Horse Society, and now forms the basis for their own electronic studbook, which it maintains using XIS systems software. At the outset of this study very little software was publicly available with which to conduct appropriate pedigree analysis. In previous investigations of a restricted dataset the Fortran software COWMATE 93 had been used to calculate rates of inbreeding (Walling, 1994). However the software GENES (Lacy, 1998) was being widely used by studbook keepers in zoos around the world and was freely available under the terms of the GUI licence. In order to facilitate genetic analysis using GENES, the data was first entered into the commercially available SPARKS (Single Population Animal Record Keeping System) software, distributed by The American Zoological Society/ International Species Information System (ISIS). The size of livestock studbooks is generally greater than any zoological collection and this was the case with the Cleveland Bay data. In order to run successfully under the SPARKS system a specially extended version was produced by ISIS, and they report that at the time, the Clevel and Bay dataset was the third largest database to run under this framework. In recent years research into Livestock Biodiversity has caught up with that conducted by wildlife conservationists, and there have been a number of useful software developments and a wider range of programmes freely available for research purposes. Of particular note are ENDOG (Gutierrez and Goyache, 2005); EVA (Berg et al., 2007); CFC (Contribution, Inbreeding [F], Co-ancestry) (Sargolzaei et al., 2006), and POPREP (Groeneveld et al., 2009). Each of these offers slightly different routines and where they have been used they are identified in the text. Pervious analysis of a partial Cleveland Bay Studbook dataset (Walling, 1994) centred around the level of inbreeding and coancestry within the breed. Calculation of an individuals inbreeding coefficient (Wright, 1922, Malcot, 1948) depends on the extent to which its ancestry is known and the number of defined generations in the animals pedigree. Whilst the absolute level of inbreeding may not be of importance, its rate of increase certainly is, being the prime parameter in assessing the additive genetic variation within the population (Groeneveld et al., 2009). The rate of inbreeding has a direct effect on one of the major statistics of breed diversity the effective population size and we will see that a variety of different assumptions and methods lead to different estimates of this. The broad spectrum of software now available has allowed detailed analysis, utilising a number of different assumptions, and each contributes to our combined knowledge of the Cleveland Bay breed. 2.2 Data Quality and Pedigree Completeness It is now accepted that the quality and validity of any genealogical analysis from studbook pedigree data is directly related to the accuracy and completeness of the dataset(Oliehoek and Bijma, 2009). Importation of the electronic studbook data into SPARKS revealed many pedigree errors, the majority of which were infinite loops, caused by incorrect identification of sire or dams. Each of these was investigated and the most likely solution found often the cause being two animals bearing the same name but different studbook number and birth-date. The necessary corrections were agreed with the Society and both studbook datasets amended, with corresponding corrections made to the paper records when the Society published its Millennium compendium edition. A number of different parameters may be used to evaluate pedigree quality. The most popular of these is that proposed by MacCluer et al, who defined a weighted index to measure the completeness of a pedigree(MacCluer et al., 1983). This index summarizes the proportion of known ancestors in each ascending generation. It quantifies the chance of detecting inbreeding in the pedigree (Sorensen et al., 2005). Calculation of Pedigree Completeness was made using PopRep (Groeneveld et al., 2009). The following formula was used to compute pedigree completeness: and Where k represents the paternal (pat) or maternal (mat) line of an individual, and ai is the proportion of known ancestors in generation i; d is the number of generations measured when calculating the pedigree completeness. Values for pedigree completeness will range from 0 to 1. Where all of the ancestors of an individual are known to some specified generation (d) then Id=1. However , where one of the parent animals is unknown Id = 0 (Groeneveld et al., 2009). 2.3 Population Demographics The software POPREP (Groeneveld et al., 2009) was used to calculate: The number of breeding males and females per year. The number of breeding animals at any one time will determine the genetic structure of the population in subsequent generations. The number of breeding females is of particular importance because the FAO and the RBST use assessment of the number of breeding females as a measure of vulnerability or endangerment of livestock breeds. ii. The Age Structure of Parents by Birth Year. Variations in breeding practice and bottleneck events may be indicated by wide variation in the age of animals being brought into or kept in reproduction. iii. Distribution of Parity of Dams at Birth of Offspring. The rate of genetic progress of a population will be influenced the turn-over of breeding stock. Where non random mating conditions exist, and animals stay in production for a long time the turn-over of breeding stock will be affected. This is particularly the case in equine breeds, where the females are non-menopausal and can remain in production well into their twenties where other age related conditions do not preclude breeding. Family Size. This relates to the number of offspring of an individual that go on to become breeding animals themselves in the next generation (Falconer and MacKay, 1996). In an idealised situation all parents have an equal chance of contributing offspring to the next generation. However, in practice this is rarely the case in production livestock. The unequal contributions both of individuals, and also the sex bias between sires and dams leads to variation in family size. A consequence of this is an increase in the rate of inbreeding and a reduction in effective population size. In calculating family size offspring were categorised into four groups All Offspring ; Selected Offspring ( going on into production); Selected Sons ; Selected Daughters (Groeneveld et al., 2009). 2.4 Average Generation Interval The generation interval (GI) is one of the most influential factors affecting the rate of genetic progress over time. In its most simple form GI is assessed from the average age of males and females in the studbook. However it is more usefully defined as the average age of the parent animals at the time of birth of their offspring, where an offspring is an animal that subsequently goes on to have progeny of its own in the studbook (Falconer and MacKay, 1996). Transfer of genes from parents to offspring will occur through four pathways sire to son; sire to daughter; dam to son and dam to daughter. The generation interval is calculated for each of these four pathways and the results averaged for each year group using PopRep (Groeneveld et al., 2009) 2.5 Inbreeding Analysis The accumulation of inbreeding within the Cleveland Bay breed has previously been reported (Walling, 1994) and has also been a topic of debate at subsequent breed conferences. Inbreeding is the mating of related individuals, which results in some loci bearing alleles that are identical by descent. This occurs because alleles from a common ancestor, appearing in both the maternal and paternal sides of a pedigree, pass though multiple offspring. The proportion of genes that are identical by descent are designated as the Inbreeding Coefficient (Wright, 1922). The Inbreeding Coefficient is defined as the probability that an individual has two identical alleles by descent. It is calculated from the formula: Where n is the number of animals in any pathway connecting the two parents of animal X back through a common ancestor, and where Fa is the inbreeding of the common ancestor. Inbreeding coefficients for each individual animal were calculated using GENES, Endog, EVA and PopRep with a high consistency of results across the range of programmes. The unavoidable mating of related animals in closed populations such as the Cleveland bay will lead to accumulation of inbreeding and loss of genetic diversity (Falconer and MacKay, 1996). This loss of heterozygosity is often expressed as an accumulation of homozygosity. The Increase in Inbreeding or Rate of Inbreeding (F), was calculated for each generation, by means of the formula where Ft and Ft-1 are the average inbreeding of offspring and their parents respectively (Falconer and MacKay, 1996). The Average Relatedness Coefficient (AR) of an individual is the probability that two alleles at a given locus belonging to two different individuals, taken at random, are identical by descent (De Braekeleer et al., 1996).. It equates to twice the mean coancestry between any individual and all of the other animals in the reference population, including the individual under consideration. The Average Relatedness of a founder is a good measure of its genetic contribution to the entire pedigree(Cervantes et al., 2008). Average Relatedness was calculated from the formula c = (1/n) 1A. Where c represents the relatedness of the individual to every animal in the population, including itself, and A is the numerator relationship matrix size nxn (Gutierrez and Goyache, 2005). Average Relatedness was calculated using ENDOG 4.6 (Gutierrez and Goyache, 2005). 2.6 Effective Population Size The Effective Population Size (Ne) is the number of breeding animals in an idealized population, that would lead to the same rate of calculated or observed inbreeding (F), as observed in the real population(Falconer and MacKay, 1996). It is a measure of diversity within a population and can be calculated in a number of different ways., principally based on either the rate of inbreeding, or on the number of parents (Falconer and MacKay, 1996). The Effective Population Size from the rate of inbreeding is computed using the classic equation Where the rate of inbreeding per generation is calculated using Ft and Ft-1 being the average inbreeding of offspring and parents respectively(Falconer and MacKay, 1996). The Effective Population Size from the number of parents is computed as Where Nm and Nf are the number of male and female parents respectively (Falconer and MacKay, 1996). This method makes the assumption that the ratio of breeding males to breeding females is 1:1, and that all individuals have an equal opportunity to contribute to the next generation. This is seldom the case in livestock populations and the tendency is for this method to overestimate effective population size by a considerable margin (Groeneveld et al., 2009). Calculations of effective population size were made using both Endog 4.6 (Gutierrez and Goyache, 2005)and POPREP (Groeneveld et al., 2009). 2.7 Founder and Ancestor Representation Studbook information was analysed in order to identify stallion and dam lines, defined respectively as unbroken descent through male or female animals only from an ancestor to a descendant(Cunningham et al., 2001). The results of this analysis were compared with those previously reported (Emmerson, 1984) (Walling, 1994). Detailed founder and ancestor analysis was carried out using Endog 4.6 (Gutierrez and Goyache, 2005), to determine: The Number of Founders, where a founder is defined as an animal that has contributed by breeding to the living population, but there is no record in the studbook of its own parent animals. Calculations were made of founder contributions to each animal in the Reference Population and of the number of living descendants of each founder. All animals with two unknown parents are regarded as founders for this analysis. In the situation where one of the parent animals is unknown this animal is also considered to be a founder. In many studbooks the analysis of the total number of founders can be misleading because many animals are recorded as sire or dam unknown, where the animal is otherwise present in the data. This is almost certainly so in the case in the recording of many dams in Cleveland Bay Horse pedigrees. In addition, some founders have been used more intensely and therefore contribute more to the current population than other founders. The effective number of founders, e, has been designed to correct for this second shortcoming(Gutierrez and Goyache, 2005). Effective Number of Founders (e) defined as the number of equally contributing founders that would be expected to produce the same genetic diversity as in the population under study (Lacy, 1989). This is computed from the genetic contributions of the Nf founders: Where qi is the genetic contribution of the ith founder to the reference and f the real number of founders(Sorensen et al., 2005). In a scenario where every founder makes an equal contribution, the effective number of founders will equal the actual number of founders. However, it is far more usual for founders to contribute unequally, and then the effective number of founders will be smaller than the true number. The genetic contributions will have converged after 5 to 7 generations (Bijma and Woolliams, 1999). Once this convergence occurs the effective number of founders will have limited usefulness as it will remain constant irrespective of later changes in the population. Pedigrees of more than 7 generations can be characterized with a high effective number of founders even after a severe, recent bottleneck. Whilst the effective number of founders is not a perfect measure of genetic diversity, it does form a basis for comparison of the effective population size and the effective number of ancestors which we qualify in the following paragraph. In a population with minimum inbreeding, the effective number of founders would be expected to be one-half the effective population size. Where the effective number of founders differs from this, there is evidence that the breeding structure has been changed since the founder generation (Sorensen et al., 2005). The Effective Number of Ancestors (a) supplements the effective number of founders (Boichard et al., 1997). It is calculated from the genetic contributions of ancestors with the largest marginal genetic contributions. Whilst genetic contributions of founders are independent and sum to one, this is not the case for genetic contributions of ancestors. The dam of a highly used sire has 50% contribution of her son, because the same genes are represented in both generations. To deal with this imbalance Boichard et al. introduced the concept of marginal contributions (Boichard et al., 1997). The ancestors contributing most to the reference population are considered, each in turn, in a recursive process. With each iteration the ancestor with the highest contribution is chosen, and the contributions of all others are calculated conditional on the contribution of the chosen ancestor. The marginal contribution thus takes into account the prior genetic contribution of ancestors already included in the recursive calculations. When summed together the product of all of the marginal contributions will b e 1. Ancestors will have a large marginal contribution to the reference population when their genes have passed through a large number of descendants, for example a sire from which the progeny has a large number of offspring themselves. (Sorensen et al., 2005). The effective number of ancestors helps to account for the losses of genetic variability produced by the unbalanced use of reproductive individuals, which is the norm in domestic equines, and also takes into account bottlenecks in the pedigree. The parameter fa is computed as Click to view the MathML source where qj is the marginal contribution of an ancestor j. The ratio of the effective number of founders and the effective number of ancestors gives an indication of the significance of any bottleneck events in population development. When the two are close to equal the population will have been relatively stable and balanced in contributions. However where the effective number of founders substantially exceeds the effective number of ancestors there is a high probability that bottleneck events have played a significant part in population development(Sorensen et al., 2005). The Effective Number of Founder Genomes (g) was proposed by Lacy (1989) to account for unequal founder contributions, random loss of alleles caused by genetic drift and for bottleneck events. It is computed by the equation: Where pi is the expected proportional genetic contribution of a founder i ; ri is the expected proportion of founder is alleles that remain in the current population, and c is the total number of contributing founders(Lacy, 1989). This gives an indication of the number of equally contributing founders with no loss of founder alleles, that would produce the same amount of diversity as found in the reference population (Lacy, 1995). The effective number of founder genome will be smaller than both the effective number of founders and the effective number of ancestors, even under minimum inbreeding, and will also be less than half of the effective population size. The scale of these differences is indicative of the degree of random loss of alleles. Alleles will be lost with every generation of a pedigree and thus the number of founder genomes will decrease as the depth of pedigree increases (Sorensen et al., 2005). 2.8 Largest Genetic Contributions The genetic contribution of an ancestor is the average genetic relationship of that ancestor with its descendents in a later generation (Woolliams and Thompson, 1994). The programme CFC (Sargolzaei et al., 2006) was used to assess the ancestors making the greatest contribution to the reference population. In addition, in order to identify unbalanced contributions of male and female animals to the Cleveland Bay Studbook, the programme POPREP (Groeneveld et al., 2009) was used to identify the Dams and Sires with the most progeny in the population; the number of progeny per Dam or Sire and also the Dams and Sires with the greatest number of progeny selected (going on to produce offspring themselves). 2.9 Population Structure Genetic structure of a population can be assessed using F- statistics(Wright, 1978). F (fixation) statistics extend the study of inbreeding coefficients in the case of sub-divided populations and consist of three parameters. FIT is inbreeding of an individual relative to the total population.FIS is the inbreeding of an individual relative to its own subpopulation. FST is the average inbreeding of the subpopulation relative to the whole population. The three indices are obtained as:- , and where and are, respectively, the mean coancestry and the inbreeding coefficient for the entire metapopulation, and, the average coancestry for the subpopulation, so that (1 FIT ) = ( 1 FIS)( 1 FST) (Caballero and Toro, 2002). All F-stats were calculated using ENDOG 4.6. Structure between subpopulations was also calculated by use of genetic distances. Neis minimum distance (Nei, 1987) was calculated as the genetic distance between subpopulations i and j given by the equation Using the data obtained in matrix format from the previous calculation, the programme TREX (Makarenkov, 2001)was used to construct phylogenetic trees to illustrate the structure and relationships between subpopulations. The analysis of subpopulations from distance matrices assumes pre-determined subgroups. In the case of the Cleveland Bay these are based on Female Ancestry lines (Emmerson, 1984). 3. Results 3.1 Data Quality and Pedigree Completeness In October 2009 the Cleveland Bay Horse Society published the 38th Volume of its studbook. Like the centenary and millennium editions, this was a compendium of recent and all existing studbook registrations. Subsequent to the publication of Volume 38, a number of new registrations have been approved by the Council of the Cleveland Bay Horse Society, so that by February 2010, the Cleveland Bay Studbook included a total of 5757 animals, of which 2763 were male and 2552 were female. In addition 230 animals were listed in the grading register and a further 212 held overseas registrations. Society data on death date of animals relies exclusively on notification from breeders or owners and is far from complete. Thus it is difficult to determine with absolute confidence the make-up of the living population. However, after analysis showed the average generation interval for the dataset to be 10 years the studbook data was revised to reflect a reference population from the most recent complete generation (1997 to 2006) of 402 individual animals. This reference population was also selected on the basis that it was the most recent for which 100% of its members had microsatellite DNA parentage testing data on file, thus providing the opportunity of comparative genealogical and molecular analyses. The analysis of mictosatellite data is reported in Chapter 4. Pedigree Completeness The pedigree file was analysed to assess the number of fully traced generations for each individual, the maximum number of generations traced and the equivalent complete generations for each animal. The maximum number of traced generations was 39. The average pedigree completeness was assessed for each animal in the studbook, for 1 to 6 and 15 generations. Completeness assessed using POPREP for animals born between 1997 and 2006 was found to be: 1 generation deep 100%; 2 generations deep 100%; 3 generations deep 99.9%; 4 generations deep 98.6%; 5 generations deep 92.6%; 6 generations deep 83.7%. Whilst POPREP selects only animals that go on to produce progeny that continue in production themselves for completeness calculations, Endog looks at the whole studbook, and thus the completeness index for the lower generations does not reach 100% as shown below. The nature of the Cleveland Bay studbook is somewhat complicated by overlapping generations, and so assessment of pedigree completeness by year-group alone cannot be considered as the only appropriate analysis. Pedigree completeness by maximum traced generations was also assessed, and was found to decrease from 89% where maximum generations was 1, to 50% at 6 generations and under 10% at 17 generations. 3.2 Population Demographics Population demographics and breeding patterns were assessed for both the whole studbook and the reference population. Of the 402 animals in the reference population, 193 animals were male and 209 female. Of the 193 males only 38 animals were registered as having been neutered but the true total is likely to be far higher. The reference population was sired by 83 unique stallions, out of 219 unique mares. Over the three most recent breeding seasons for which complete records are available, (2005 2007), 153 animals are registered in the studbook, sired by 52 different stallions, out of 120 different mares. These latter s most closely represent the current actively breeding global population. In 2008 there were 88 male animals holding Society stallion licences. 40 of these licensed animals have progeny registered in the studbook. 5 shows the pattern of annual registrations in the studbook. Registrations prior to 1885 will have been retrospective, with the ones prior to around 1830 based on annecdotal evidence or family breeding records, as they will have predated the living memory of the vast majority of early Society members and breeders. The peak between 1885 and 1900 will probably reflects the enthusiasm brought about by the newly formed CBHS, which was clearly already in decline before the start of World War 1 in 1914. In the interwar years annual registrations seldom reached double s. Between 1945 and 1960 it decreases further, such that the breed goes through a genetic bottleneck that has previously been identified(Walling, 1994). s 6 and 7 show the number of animals in active reproduction, listed by the year of their birth, with peaks corresponding to the pattern of registrations already described. 7 reveals that of the stock being produced only about 33% of females go on into reproduction themselves whereas the for males is closer to 50%. This may be masked by the practice of some breeders not registering male foals that are not intended to be kept entire, but go on to be neutered and sold on in the past as working animals or in more recent years for sport and leisure purposes. European Union legislation, brought into force in the late 1990s now reuires every equine to have passport documentation. This will have brought to a halt the practice of non registration of neutered animals, although not all of this group will continue to be registered through the CBHS (for social or political reasons), with other general registry bodies being available. 8 focuses onto the pattern of breeding since 1950 and shows the accute nature of the bottleneck, with no male progeny being produced or selected between 1952 and 1954, at a time when there were only 4 registered stallions. The number of offspring being produced shows a significant increase in the early 1970s, possibly associated with the enhanced general awareness of rare breeds brought about by the founding of the Rare Breeds Survival Trust, and the identification of the endangered status of the Cleveland Bay horse. The Rare Breeds Survival Trust (RBST) maintains a Watch List of endangered animals, in which Category 1 Critical is the most endangered status. Three equine breeds are listed in this category which includes the Cleveland Bay Horse, alongside the Suffolk Punch and the Eriskay Pony. To be in this list a breed has to have fewer than 300 breeding females. To asses this the RBST take a breeding female to be a female who has produced a registered foal. To calculate the number of adult breeding females for the watch-list, the trust uses the number of female registrations averaged over the last three full years and applies a multiplier (6.67) to give an estimate of the number of adult breeding females. By adopting this system females over a certain age which may still have produced registered progeny are not discounted from the analysis. Using the data provided by the CBHS (late in 2009 for the 2010 RBST Watch-list) the above calculation gives an average of 24 female registrations per year over 2006-2008, to which the multiplier of 6.67 is applied, resulting in an estimation of 162 breeding females. Whilst there has been an increase in the number of breeding females since the early 1970s, with the calculated numbers approaching the 300 in the mid 80s and again in the 90s, the recent trend is for a decline in numbers. The most recent of 162 breeding females indicates that a doubling of the breeding stock would be required for the breed to be downgraded from its present Critical status. 10 shows that the mean age of reproduction for both males and females is at 5 years old. The studbook shows some animals in reproduction at 1 year of age, but this is certainly an anomaly caused by retrospective registration practices at the formation of the studbook, and an attempt to remove loops from pedigrees, with animals having recorded birth dates before their parents. Whilst some of the records of 2 year old animals in production may also be erroneous, it has certainly been a practice amongst some Cleveland Bay breeders to use young entire males to cover their mares, and this practice will no doubt be of benefit in balancing contributions of overused stallions. From its peak at 5 years old the number of females in production decreases steadily through to 15 years. There are certainly a number of mares going on to produce progeny well into their twenties and these are seen clustered together in the graph as 16+. Similarly the pattern of males in reproduction decreases from 5 years of age but also extends beyond 16 years of age, with recent records in the studbook of one rediscovered stallion being successfully brough back into reproduction by natural covering at 29 years of age. s 11 and 12 illustrate the parity of females as best as can be done, with no records of service being available. Using the number of foals registered may not show the true pattern as animals being lost before full term gestation and during or soon after birth will not be included in the data. Family Size The breeding pattern revaled by this analysis shows that even whilst the breed was going through a bottleneck during the 1950s and 60s some diehard breeders were successfully getting 6 or more pure bred foals from some mares. Whilst well over 200 mares have successfully bred 5 pure bred foals there are a significant number producing 10 or more, with one mare being recorded as having 15 pure bred foals. Some of the mares included in the analysis will no doubt have been bred to non-Cleveland Bay stallions at some time during their breeding life, and these foalings will not be reflected in the tables. This practice of non pure, or part-bred breeding has been in existence throughout the history of the Society, with considerable demand for Cleveland Bay x Thoroughbred animals as coaching horses prior to the mechanisation of transport, and in more recent years as sport and competition horses. 13 shows the comparative analysis of number of progeny per sire. Whilst over 250 individuals are recorded as having sired only one progeny, and 125 having only two, a significant number of stallions are recorded as having over 20 progeny. A small number of stallions are recorded as having over 60 progeny , with the most prolific animal having in excess of 250 foals. This unbalanced pattern of breeding is of importance in explaining the loss of diversity in the breed and is explored in more detal later in this chapter,in the analysis of Genetic Contributions. The influence of parent animals to the subsequent generations can only be properly shown by analysis of their progeny that go on into production, and have live offspring themselves. s 14 and 15 illustrate the number of selected offspring being bred by dams and sires respectively. 16 shows the overall pattern of breeding since 1900. By selecting only the post 1900 data, the erratic breeding pattern of the early Society years is ignored, and a significant increase in the number of progeny per sire in the post bottleneck period is revealed. This trend peaks again in the mid 1960s and late 70s, then begins to decrease. However the graph shows that there is still a very unbalanced breeding pattern, with individual animals, particularly males, making large contributions to subsequent generations. 3.3 Average Generation Interval The average generation interval for each breeding year was found to range between 5.5 and 13 years, being at a minimum in the immediate post WW2 period 1946 to 1950, which coincides with the previously identified genetic bottleneck(Walling, 1994). The average generation interval for the whole population comprises four component parent progeny pathways, and when the separate components are examined, wide variations are evident, particularly in the sire-son pathway, with a maximum of 25 years in 1904 and a of 19.9 years as recently as 2003. s 18 and 19 illustrate the generation interval by each of the four selection pathways in both the whole and reference polulations. The average generation interval for both populations is 10 years which is commensurate with that reported for other equine breeds(Zechner et al., 2002, Hamann and Distl, 2008). 3.4 Inbreeding Analysis Calculations show mean inbreeding across the whole population of 7.8% with an associated mean average relatedness of 8.3%. The graph shows a near linear trend in accumulation of average inbreeding between 1885 and 1985, by which time it was approximately 20%. Since 1985 the rate of accumulation has slowed, with an average in the reference population of approximately 21%. The pattern of inbreeding in the Reference Population, over the period 1997 to 2006 is shown in Table 1 (source POPREP). YEAR No Animals Min Inbreeding Max Inbreeding Average Std 1997 57 0.1327 0.2943 0.2072 0.0305 1998 46 0.1540 0.2943 0.2139 0.0254 1999 54 0.1448 0.3156 0.2126 0.0317 2000 64 0.1654 0.3079 0.2139 0.0269 2001 37 0.1783 0.3132 0.2186 0.0250 2002 46 0.1830 0.3084 0.2218 0.0263 2003 52 0.1830 0.3017 0.2173 0.0227 2004 63 0.1629 0.2852 0.2133 0.0231 2005 54 0.1100 0.2580 0.2102 0.0219 2006 76 0.0925 0.2616 0.2065 0.0253 Table 1: Inbreeding Coefficients F of Reference Population by birth year 21 shows the rate of change of both the Inbreeding Coefficient and the Additive Genetic Relationship between 1901 and 2009. The rate of change of the average inbreeding coefficient, based on slope regression was 0.00214, which represents a F per generation of 0.02709.The rate of change in the Average Genetic Relationship (also based on slope regression) was calculated as 0.00202 per year. This results in a f per generation of 0.02629. From these results the average Effective Population size for the Cleveland Bay Horse breed over the period 1901 to 2009, based on f was 19 and from F 18 (source POPREP). Significant difference between the inbreeding coefficient and average genetic relationship may be masked by overlapping generations, and to identify any such trend Table 2 presents the data by complete generations. Complete Generations N F AR NE 0 1562 0 0.01508601 1 1127 0.0109910 0.04204081 45.4 2 532 0.0409840 0.06222444 16.4 3 202 0.0733043 0.1084541 14.8 4 304 0.1299420 0.1471427 8.1 5 605 0.1761405 0.1711177 9.4 6 694 0.1931363 0.1795912 24.2 7 342 0.2047071 0.1813364 34.8 8 54 0.2167604 0.1838901 32.9 Table2: Change in inbreeding coefficient and average relatedness for 9 fully traced generations. (Source ENDOG 4.6) Note that with and average inbreeding of 0 for animals of no traced generations it is not possible to calculate the Effective Population size from the rate of inbreeding. 21 shows the discrepancy between Average Genetic relationship and Inbreeding across a maximum of 39 generations. The scale of the difference is indicative of the deviation from random mating and unbalanced genetic contributions. 3.5 Effective Population Size: Effective population size was calculated by two different methods (rate of change of inbreeding and number of parents), both of which are illustrated in 22. This illustrates the wide variation in Effective Population size brought about by the fluctuations in inbreeding at the time of the start of the studbook, during two world wars and the post war bottleneck. The Effective Population Size calculated from the number of parents appears to be more stable, with two peaks at about Ne = 100. The first occurs in about 1888, shortly after the founding of the studbook. The second occurs in more recent times, approaching a maximum of Ne = 105 in 2006 as shown in Table 3. 3.6 Founder and Ancestor representation Founders Gene drop analysis on the whole population using GENES (Lacy, 1998) identified 182 founders with a mean retention of 0.033. The number of founder genomes surviving was 6.015 and the number of Founder Genome Equivalents 2.219. The fraction of source diversity retained was 77.5% with a corresponding source diversity loss of 22.5%. The proportion of ancestry known was 0.330 reflecting the fact that in early volumes of the studbook it was often the case that only a record of the sire of an individual animal was made, with its dam recorded as unknown. Effective Number of Founders / Ancestors For analysis of The Effective Number of Founders/Ancestors (Boichard et al., 1997) the base population was taken as the animals with both parents known. This was found to be 4947 animals, and was substantially smaller than the number of individuals considered in other whole population analyses. The number of ancestors contributing to this base population was 614. For the base population, the Effective Number of Founders was found to be 77 and the Effective Number of Ancestors 24. 8 Ancestors explain 50% of the genetic make-up of the base population. Analysis showed that only 30 Ancestors contribute to the 402 individuals in the Reference Population (1997 to 2006). In the Reference Population the Effective Number of Founders was reduced to 61 and the Effective Number of Ancestors was down to 9. 50% of the makeup of Reference Population was explained by only 3 Ancestors . Ancestors were selected following Boichard et al. (1997), while founders were selected by their individual Average Relatedness coefficient (AR(Gutierrez and Goyache, 2005)). Effective Number of Founder Genomes Calculations on the base population using CFC show the founder genome equivalent (Lacy, 1989)to be 2.366.. Sire Lines A total of 11 stallion lines were identified in the pedigree. However, as previously reported (Walling, 1994), only one paternal ancestry line is present in the living population. This line had previously been traced back and attributed to Wonderful Lad (SB No 361), but more detailed analysis, made possible by complete digitisation of the Cleveland Bay Studbook, traces this line a further five generations to the stallion Skyrocket (SB No280) born in 1805. Further research of sources outside the Cleveland Bay Studbook indicates that this animal is probably descended in the male tail line from the thoroughbred founder Byerley Turk. The male pathway for this is Byerley Turk Jigg Crofts Partner Tartar Herod Highflyer Skyrocket (TB) Skyrocket (CB). The pedigree of Highflyer is illustrated in 23 and shows that all three of the imported horses said to be founders of the thoroughbred (Byerley Turk / Darley Arabian and Godolphin Arabian) are present to some degree in the pedigree of the Cleveland Bay founder Skyrocket (Studbook No.280). In his study Walling linked Wonderful Lad 361 to what were identified as 4 bottleneck founders(Walling, 1994). In fact these are not strictly founders but are more appropriately described as ancestors (Boichard et al., 1997). 25 shows the pathway from the founder Skyrocket to Wallings 4 Bottleneck Founders. Also highlighted are the two stallions, Newton 216 and Reform 653, that Emmerson has identified as founders of male lines 1a and 1b. There is a single male pathway back to Skyrocket from the common sire of these two animals (Sportsman 299). Dam Lines Analysis of the female members of the studbook identified a total of 17dam lines (Table 5). As previously reported (Emmerson, 1984, Walling, 1994) only nine maternal ancestry lines are present in the reference population. Three of these lines (2, 4 9) are only represented in the living population, in direct female descent, by one or two individual animals. The three most common of the maternal lines (1, 5 6) make up 70% of the present female population. Interestingly, analysis of the relative contributions of the most influential maternal ancestry lines to the genome of the reference population reveals that some of the lines least well represented in direct descent actually continue to make a substantial genetic contribution as shown in Table 6. Table 5: Female Tail Lines / Founders Maternal Line Contribution to Reference Population TWO Depper 39 27.44% ONE Stainthorpes Star 25.92% SIX Trimmer 268 21.54% FIVE Depper 42 15.89% THREE Dais(y) 318 2.13% FOUR Marvellous 72 1.29% EIGHT Church House Queenie 0.26% Table 6: Contribution of Female Founders to Reference Population Of the 2793 female registrations some 1271 animals fall outside the 17 female lines. This can be explained by the fact that in the early days of the stud book many horses only had their sires recorded. In comparison of the 2831 registered males, only 176 do not belong to one of the eleven male lines. Genetic Contributions The relative genetic contributions of each of the female tail lines to the whole and reference populations are shown in Table 7. Maternal Line N Whole Pop % Evol Rate Of Whole Pop N Rep Pop % Evol Rate Of Ref Pop 1 424 7.819992623 11.38% 77 19.15422886 25.93% 2 231 4.260420509 11.59% 3 0.746268657 27.45% 3 269 4.961268904 0.52% 56 13.93034826 2.14% 4 26 0.47952785 0.08% 0 0 1.29% 5 500 9.221689414 9.32% 82 20.39800995 15.89% 6 611 11.26890446 o% 115 28.60696517 21.54% 7 177 3.264478052 0.00% 61 15.17412935 8 78 1.438583549 0.18% 4 0.995024876 0.27% 9 3 0.055330136 0.01% 0 0 0 3103 57.2298045 4 0.995024876 Table 7: Relative contributions of maternal ancestry lines to the evolution of the whole and reference (1997- 2006) populations. Note that the population sizes reported here include both male and female representatives of the Maternal Lines, so N is higher than shown in table 4 where only female members are shown. The reason behind this is that when considering a female tail line some of the genetic material (in particular mitochondrial DNA) can only be passed down the female to female pathway. Thus whilst a son will carry its dams mitochondrial DNA it will not pass it on to its own progeny. s 26 and 27 show the 30 sires with most progeny and most selected progeny (those that go on to reproduce themselves) in the whole population. Whether the stallion Prince George M235 actually produced 285 progeny is debateable. As he was a fashionable stallion at the time his progeny would have been highly marketable and sold for a premium. Whilst not questioning the integrity of breeders of the day, it is not improbable that some animals were registered as by him that were otherwise bred and without the modern day parentage testing and registration requirements the truth may never be revealed. However, there is more certainty about the prolific use of certain modern day stallions, particularly Forest Superman M1925 and Storth House Temptation M2054. In the full version of his dissertation Walling comments on the problems of overuse of stallions like Storth House Temptation. This possibly sensitive observation was omitted when reported in the Cleveland Bay Horse Society Studbook Vol 33.(Walling, 1994), the stallion in question still being alive and in use at the time. 28 illustrates the 30 most prolific dams in the whole population, whilst 29 shows them ranked by the number of progeny that go on into reproduction. Interestingly both Depper 42 and Dais 318 feature in this list. They are the founding mares of line 5 and 3 respectively. Founders Table 8 details the 10 founders making the greatest contribution to the reference population as evaluated by Endog. In making these calculations the base Population (one or more parents unknown) was assessed at 1117 animals, and the Actual Base Population (one unknown parent = half founder) as 655. The effective population size of founders was assessed to be 101.00. Name Studbook Number Sex Year of birth Contrib (AR) v1 v2 Festivity FSB Female ~1877 0.0416382 0.0082554 0.00845627 Bilsdale Violet 1019 Female ~1889 0.0108296 0.0146294 0.00151867 Emulator 103 Male 1844 0.00993098 0.00209102 0.00213754 Cleveland Champion 458 Male 1810 0.00772717 0.00154109 0.00157881 Salesman 271 Male 1846 0.00699245 0.00149669 0.0152981 May 73 Female 1878 0.00578839 0.00109138 0.00111974 Park Polly 843 Female 1876 0.00578839 0.00109138 0.00111974 Barley Harvest 447 Male 1796 0.00170209 0.000413771 0.000421677 Skyrocket 280 Male 1805 0.00132344 0.000299756 0.000305968 Sportsman 430 Male 1860 0.000729633 0.000123869 0.000128294 Table 8: The 10 founders contributing the most to the reference population. V1 = Contributions of genes of founders to the average inbreeding coefficient V2 = Contributions of genes of founders to the average coancestry Ancestors Ancestors were selected following Boichard et al. (1997). Table 10 sets out the male ancestral contributions to the reference population assessed using CFC(Sargolzaei et al., 2006). Of the four modern day founders identified by Walling only three appear as ancestors in the table. The most significant of these is Lord Fairfax M1875, who makes the greatest contribution of just below 22%. Also appearing in the rankings are Apollo M1857 at 11.23% and Cholderton Druid M1859 ranking 31st with a contribution of 6.50%. Interestingly Wallings other modern day founder Cholderton Cockade M 1858 does not appear in this table. However, also of note is that a number of pre-bottleneck stallions make a significant contribution to the reference population, including Morning Star (17.03%); Aislaby Lad (16.20%) and Cholderton Ryecroft (15.35%). Table 9: Male ancestral contributions to the reference population Table 10: Female ancestral contributions to the reference population Table 10 sets out the corresponding largest female contributions. Of note is the fact that a number of these ancestors fall outside the 17 identified female ancestry lines including Star of Hope and her daughter Star of the Sea, Woodland Starlight and the mare Beauty who sits much earlier in the studbook. These proportional contributions are shown by ancestry line in 30. Note that the sum of genetic contributions of ancestors can be 1 as there may be sharing of genes with other ancestors. 3.7 Population Structure Within population genetic differentiation was assessed using F-statistics (Wright, 1931, Weir and Cockerham, 1984) computed from the pedigree subdivided by female ancestry line. Fst reached 0.039427; Fis = 0.003891 ; and Fit = 0.035689. This indicates a significant differentiation of the population at this level and a departure from Hardy-Weinberg equilibrium. In order to visualise this differentiation and the possible relationship between the different lines two different paired distance matrices were constructed. The first was based on Fis-Fst relationships and the second standard distance(Nei, 1973). This data was input into the web based version of the programme T-Rex (Makarenkov, 2001) to produce visual phylogetetic trees ( s 31 to 34). A variety of tree drawing methods were implemented, with consistency between methods regarding both the number of sub populations at the root of the tree and the identity of those populations. The diagrams show that lines 3, 1 and 5 consistently appear at the root of three clusters or clades in each of the trees. Discussion The overall results presented here highlight the significant losses of founder representation that have occurred in the Cleveland Bay Horse population. Approximately 91% of the stallion and 48% of the dam lines are lost in the reference population. The unbalanced representation of the founders is illustrated by the effective number of founder animals (fe) and the effective number of ancestors (fa). The parameter fe constitutes over a third of the equivalent number of founder animals for the RP, whilst the ratio fa/fe is 22.5%. This ratio is substantially lower than that reported in other horse breeds such as 41.7% in the Andalusian (Valera et al., 2005), 54.4% in the Lipizzan (Zechner et al., 2002) and 38.2% for the endangered Catalonian donkey (Gutierrez et al., 2005) The average inbreeding computed for the Cleveland Bay Horse at 20.64%in the RP is substantially higher than most of the values reported in the literature (Valera et al., 2005), with typical values ranging from 6.5% to 12.5%. Although most of these inbreeding values have been computed in breeds with deep pedigrees such as Andalusian, Lipizzan or Thoroughbred there are significant differences in population sizes, and the accumulation of inbreeding in populations of restricted size will occur at a greater rate. The smaller the number of individuals in a randomly mating breed the greater will be the accumulation of inbreeding because of the restricted choice of mates. The Cleveland Bay horse is therefore predisposed to inbreeding and associated loss of genetic variation. In the reference population of 402 individuals the Effective Population Size (Ne) computed via individual increase in inbreeding was 27.84. The Effective Population Size (Ne) computed via regression on equivalent generations was 26.29. Inbreeding and genetic loss under random mating will occur at 1/2Ne per generation. Thus in the Reference Population where Mean Ne is 32.32, under random mating we can expect inbreeding to accumulate at 1.5% per generation. It is generally accepted that livestock breeds should be managed to keep the accumulation of inbreeding below 1% per generation. The results obtained in this investigation are a clear indication of the need for proactive breed management in order to maintain founder representation and maximise genetic diversity. This is reflected by the genealogical FIS values. This parameter characterises the mating policy meaning the departure from random mating as a deviation from Hardy-Weinberg proportions. Positive FIS values mean that the average F value within a population exceeds the between individuals coancestry, thus indicating that matings between relatives have taken place(Caballero and Toro, 2000, Gutierrez et al., 2005) Moreover, the average AR values computed for nine complete generation shown in Table2 are approximately equal to the value of F. In an ideal scenario with random matings and no population subdivision, AR would be approximately twice the F value of the next generation.(Goyache et al., 2003) However until the year 2002, F has continued to increase in the Cleveland Bay Horse population. Departure from random mating will have been influenced by a number of factors common to restricted populations of domesticated equines. These include selection by breeders for particular lines of descent; natural differences in fertility between individuals; a restricted number of male animals leaving significantly more offspring than females and geographic distribution of animals and breeders making some matings far easier than others. This is indicative of the typical practice of the larger studs, where breeding tends to be done in pasture by live cover, with stallions running with the mares. Often only one stallion is used per year per herd and the same stallion may be retained for several breeding years. This practice is compounded by breeders with only a small number of breeding females sending their animals to run with these herds or to be covered in hand by the same stallion. This pattern of stallion use has different implications for the genetic diversity of the Cleveland Bay Horse compared to the alternative of mares travelling to stud to be covered in hand by a wider range of stallions, that do not have their own herds of mares(Luis et al., 2007). Although this latter practice has clear benefits in conservation programmes, there is the danger of inappropriate matings bringing the more common and less frequent alleles together. Whilst such matings increase the frequency of the rarer alleles they simultaneously increase the frequency of the more common ones(Lacy, 2000), highlighting the need for in depth understanding of the genetic diversity of any rare breed and for an effective management plan for its maintenance. What is evident from this analysis is that inbreeding has accumulated at an undesirable rate in the Cleveland Bay breed. In addition there has been a significant loss of founder representation in the living population. This loss of diversity is reflected in the low effective population size for the breed. Taken in combination this provides substantial evidence that the uncoordinated random breeding practice that has taken place in the past is not the best strategy for the preservation or genetic health of the breed. In the next chapter we shall examine what can be understood about the genetic diversity of the breed at molecular level, and learn whether the use of microsatellite DNA analysis can shed further light on within-breed population structure. References. BERG, P., SORENSEN, M. K. NIELSEN, J. 2007. EVA Interface Program. University of Aarhus. BIJMA, P. WOOLLIAMS, J. A. 1999. Prediction of Genetic Contributions and Generation Intervals in Populations With Overlapping Generations Under Selection. Genetics, 151, 1197-1210. BOICHARD, D., MAIGNEL, L. VERRIER, E. 1997. The value of using probabilities of gene origin to measure genetic variability in a population. Genet. Sel. Evol. , 29, 5-23. CABALLERO, A. TORO, M. A. 2000. Interrelations between effective population size and other pedigree tools for the management of conserved populations. Genet Res, 75, 331-43. CABALLERO, A. TORO, M. A. 2002. Analysis of genetic diversity for the management of conserved subdivided populations. Conservation Genetics, V3, 289. CERVANTES, I., MOLINA, A., GOYACHE, F., GUTIERREZ, J. P. VALERA, M. 2008. Population history and genetic variability in the Spanish Arab Horse assessed via pedigree analysis. Livestock Science, 113, 24-33. CUNNINGHAM, E. P., DOOLEY, J. J., SPLAN, R. K. BRADLEY, D. G. 2001. Microsatellite diversity, pedigree relatedness and the contributions of founder lineages to thoroughbred horses. Animal Genetics, 32, 360-364. DE BRAEKELEER, M., DAIGNEAULT, J., ALLARD, C., SIMARD, F. AUBIN, G. 1996. Genealogy and geographical distribution of CFTR mutations in Saguenay Lac-Saint-Jean (Quebec, Canada). Annals of Human Biology, 23, 345. DUCRO, B., BOVENHUIS, H., NEUTEBOOM, M. HELLINGA, I. 2006. Genetic diversity in the Dutch Friesian horse. Proceedings of the 8th World Congress on Genetics Applied to Livestock Production: 13-18 August 2006. EMMERSON, S. 1984. Cleveland Bay Horse Society Centenary Studbook, Cleveland Bay Horse Society. FALCONER, D. S. MACKAY, T. F. C. 1996. Introduction to Quantitative Genetics, Essex, England., Longman. GLAZEWSKA, I. JEZIERSKI, T. 2004. Pedigree analysis of Polish Arabian horses based on founder contributions. Livestock Production Science, 90, 293. GOYACHE, F., GUTIERREZ, J. P., FERNANDEZ, I., GOMEZ, E., ALVAREZ, I., DIEZ, J. ROYO, L. J. 2003. Using pedigree information to monitor genetic variability of endangered populations: the Xalda sheep breed of Asturias as an example. Journal of Animal Breeding and Genetics, 120, 95-105. GROENEVELD, E., WESTHUIZEN, B., MAIWASHE, A., VOORDEWIND, F. FERRAZ, J. 2009. POPREP: a generic report for population management. Genet. Mol. Res., 8, 1158-1178. GUTIERREZ, J. P. GOYACHE, F. 2005. 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The Controversy Of Laramie, Wyoming - 1735 Words

Laramie, Wyoming is known as a mostly conservative, republican party majority, which is why there were no hate crime laws that had to deal with sexual orientation or gender identity. Republicans typically were Christians meaning they were against homosexuality and believed it was very wrong. The Laramie officials did not pass any law for about ten years, which showed the Laramie community that their own government did not approve of homosexuals. The government in Laramie is a major influence on a citizen’s political opinion and how they feel on certain topics like homosexuality. If the government does not show approval, how will the citizens of Laramie gain a major acceptance towards homosexuals? Even the governor of Wyoming did not†¦show more content†¦This bill â€Å"was called Resolution Seventeen, a bill defining marriage in Wyoming as being exclusively between a man and woman† (Kaufman 163). Connolly passionately fought against such a piece of legislation arguing homosexuals should be able to be able to marry as well. This did not go as well since the Laramie legislation consisted of mainly conservative republicans refusing to allow anything, but the traditional style of marriage. Republicans argue, †since the beginning of civilization, in every known society, governments have recognized a marriage between a man and a woman because it provides the next generation outstanding citizens and is the only means of melding two sexes into a stronger and more complete whole† (Kaufman 164). This political party is very strict to their views and use history to show how a traditional marriage is the only logical way to continue a working society. Republican believe since homosexuals are unable to reproduce, they are unwilling to have a functional family, which is why they do not agree with same-sex marriage. Since the republicans did not pass this bill same-sex marriage is still currently today illegal in Wyoming. By not passing this bill in Laramie this shows that there is still ignorance throughout both the town of Laramie, the state of Wyoming, and the country of the United States of America. This ignorance will continue to roam throughout Laramie until change happens fully accepting all homosexuals as equal members of theShow MoreRelatedConservatism, By Moises Kaufman And The Tectonic Theater Project1722 Words   |  7 Pagesthose who oppose it to preserve a detestable status quo. The play The Laramie Project by Moises Kaufman and the Tectonic Theater Project introdu ces some of those people who live in the quiet town of Laramie, Wyoming. It has been 18 years since the brutal murder of Matthew Shepard, a gay resident of Laramie. After that tragedy, has anything changed? One answer comes from Laramie resident Rebecca Hilliker who states in The Laramie Project: Ten Years Later by Moises Kaufman that, â€Å"On the surface thingsRead MoreHate Crimes: Society Reaction to Change Essay1431 Words   |  6 Pagesrejection? What does that tell us about our society? Personally I believe that we in society are afraid of change and things that we cannot control or understand. When things are different we tent to judge and try not to fully understand. The biggest controversy in our society today is sexual orientation of individuals, which can mean transgender and gay. What does these terms mean? When an individual is gay they have a romantic attraction, sexual attraction or behavior between someone of the same sex.Read MoreThe Regulation Of Lgbt Orientatio n2084 Words   |  9 Pageseven the most incredible scholars cannot completely comprehend the nature of the violence until they experience it for themselves, and Clementine Von Radics is one of these individuals, explaining her thoughts as she traveled through the town of Laramie, Wyoming, the sight of Matthew Sheppard’s horrendous murder, an act committed based solely on discrimination. Clementine Von Radics, within her short narrative Road Trip across the Country, elaborates on the nature of LGBT* discrimination by identifyingRead MoreNative American And Native Americans1156 Words   |  5 Pagesexperience these alterations were the Dakota and Lakota Sioux. The Dakota Sioux were located in South Dakota, Minnesota, Nebraska, Montana, and North Dakota. This tribe had to go through many changes throughout their history. 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But despite the benefits, changing the diversity mix of law enforcement agencies reflects the same challenges of doing so in other sectors of U.S. industry. One controversy focuses on the tests required to become officers. About 83% of law enforcement departments require applicants to pass a test on reading, writing, and around its use. For example, in Suffolk County, New York, where African Americans make up 4.8%

The Origin of Civilization - 1899 Words

The idea of free expression of political thought really came into its own in 18th century Europe. Writers and thinkers like Adam Smith, Rousseau, and Edmund Burke shared their ideas that still give reasons for consideration even today. Rousseau gave the underpinnings for the French Revolution. Smith gave us the foundations for modern economic theory. Burke gave us the idea of Conservatism, which fathered all other -isms. While all three of these writers gave us so much, it is important to look back and and see not only where their ideas came from, but also how there were in some ways just different interpretations of the same thing, and where they were in stern disagreement. One of the most important arguments that these men debated is†¦show more content†¦They placed a sovereign at the (to) top analogous to the father, and this sovereign is free to lead his people as he sees fit. What Rousseau does is free himself from this constraint, and opens up the possibility of totally different social structures. The idea is that the sovereign is only in charge as long as the people let him. He states this argument in chapter V of the Social Contract. Burke is the very antithesis of Rousseau. Burke was an Englishman and part of the Whig political party. Because of his political involvement, much of Burkes work sounds like a political speech. For him, a new theory could never carry as much weight as the preexisting school of thought. Old ideas are the way they are because they have stood the test of time, and this is a separate and cumulative legitimacy to that of any logic or reason there is to them. This is especially true of political organization. If the current structure has lasted for many generations, then there must be something to it. At the most it may need small changes every now and again. Burke makes this explicit when he says about English people: â€Å"We know that we have made no discoveries, and we think that no discoveries are to be made in morality, nor many in the great principles of government, nor in the ideas of liberty, which were understood long before we were born...†3 This line and the lines before it make it clear what Burke thinks of the startlingShow MoreRelatedThe Origins Of The Harappan Civilization1060 Words   |  5 PagesWhen the Harappan civilization began to decline, the Aryans filled the power vacuum. The Aryans were to become the new race of Indians, spreading throughout the entire Indian subcontinent. They witnessed the implementation of the caste system into Indian society and the development of the earliest Indian religion: Brahmanism. The caste system determined what people would do everyday and divided the population by social status, playing a huge role in the entire society. 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Public Health Policy Theoretical Framework

Question: Describe about the Public Health Policy for Theoretical Framework . Answer: Introduction: Development of a proper and justifiable theoretical framework is the first important step that a policy maker will adapt in order to design a particular public health policy. This is important in the sense that it will lead the policy makers to summarize all the important pros and cons that would be faced during the implementation thereby making the policy makers aware of the challenges and the demands that the situation requires. Moreover creating a theoretical framework also helps in the establishment of the clear motives and aims that would lead to a successful policy, including all necessary steps and actions and excluding any negative aspects that may help in the benefit of public. In such a scenario, policy making for the family and domestic violence in Australia also would require a theoretical framework that would result in holding this concerns faced by the victims and proper implementation of laws and legislatures fir successful inhibition, making people educated enough to handle situation more maturely (Anfara Martz, 2014). Theoretical framework and application: While designing the theoretical framework for the increasing family and domestic violence all over Australia, the first step would be to recognize the perpetrator and the victims and thereby understand the various kinds of violence and the dominancy that can be seen in every case. Recognizing the level and the extreme extent of violence that can occur in every of the cases, a proper monitoring system has to be developed along with proper categorization and the levels in which each of the cases has to be assigned (Dunn, 2015). A general policy for all the victims is not benefitting in the sense because it judges all the cases from the same sense of importance irrespective of the level of violence. Therefore, it is indeed importance that policies should be prepared aiming the growing complicacies of different cases (Sabatier Weible, 2014). Proper punishments or counseling (in case of complicated cases) of the perpetrators are also should be inculcated. The next important criteria is t hat proper governing Medias should be included such as proper police forces posted on emergency situations can perform their duties well, proper formation of emergency helpline in order to prevent violence and harm from perpetrators. Another important arena that should be covered during frame working the policies are the proper arrangement of the shelter homes and social communities that would act as the shelter places for the victims and also help them to come out of depression, shocks and fear (Kettner, Moroney Martin, 2016). Councilors should be appointed who would help the victims to come out from these incidences and start life afresh. Another thing that the policy can include is to provide small jobs to the victims so that they can support them financially and help them to achieve back their self-respect (Lin, 2012). This framework is believed to cover all the aspects of FDV that when implemented into a policy can be believed to bring out a successful and overall acceptance l eading to the benefit of the Australians. Proper conductance of surveys for the correct reporting of the cases is mandatory (Larkin, Felitti Andra, 2014) Identification of existing policies: Out of the various policies, that government has published in the recent years the one that has garnered attention is the The National plan to reduce Violence against women and Children that has been conducted with a plan for 2011 to 2022. Another policy introduced by department of prime minister and cabinet is also seen to cover a wide array of different aspects of family and domestic violence and treating the victims from the grass root of the cause and wide advantages for the victims who faced family violence (Department of Prime Minister and Cabinet, Australia, 2016). Family and Domestic violence strategy plan has been conducted by the department of human services with several aims to reduce FDV with a plan from 2016 to 2019. Western Australia Strategic plan has been implemented by the government of Western Australia that looks solely over the protection of children and intimate partners who had been seen to be most prone to get effected from FDV. This is introduced with a planni ng of 2009 to 2013 and has been reintroduced. Identification of Stakeholders groups: In the different policies that had been implemented in the FDV, cases a variety of stakeholders have been found to be involved. Apart from the victims starting from children, women, elders of the households etc. and the perpetrators, health care professionals, social workers arranging the communities, ministers and policy makers also take active part from the benefit of the nation and thereby reduce the frequency of violence (Department of Prime Minister and Cabinet, Australia, 2016). The police department along with ministers involved in the public health domain have shown much interest over the issue and taken active holds of the cases reported. Counselors involved in the development of mental stability of both the victim and the perpetrators appointed by the different agencies and social communities do commendable jobs, which are praiseworthy. All of them have to been found as effective stakeholders involved in active participation for making Australia a violence free nation. Identification of political contexts: From the various policies mentioned above, one can have a clear idea about the interest of the ministers in developing a violence free nation. In the policy given from the government of Western Australia , one can see how the Minister for child protection and community services namely Robyn McSweety have helped in providing strategic plans for the protection of violence over children and intimate partners gaining appreciation from the entire Western Australia helping in her political career as well (Western Australian Government, 2016). A proper establishment of a policy that would save the victims would not only provide benefit t nation but can also assure certain position of Ministers to be assured by the voters due to their success in preventing FDV. Department of Prime Minister and Cabinet as well as social services and Human services have productive ministers giving away their best approach for the proper implementation of the effective policies. Summary of the policies: The policy produced by the Department of Prime Minister and the Cabinet mainly has included their own employees working under this sector an takes care to protect their employees from FDV and also prepare safe environments for victims so that they can stabilize themselves and prepare an environment suitable for their recovery. It has included emotional, verbal, sexual, social, economic, psychological, spiritual and physical abuse that not only affects individuals at a personal level but also affects the workplace (Department of Prime Minister and Cabinet, Australia, 2016). As a result, this policy has taken steps to empower individuals with variety of financial, social and mental support from the managerial level as well as for the colleagues in every ways possible even by providing money and mobile phones on loan and keeping restrooms for them as well as for their children. Leave arrangements, advance payment of salary, flexible shifts and working arrangements, short or long-term re location of the victims to another office of PMC office are several options provided to them. The National Level to reduce Violence against Women and Children was conducted through two stages first by the introduction of First Action plan that involved the foundation of a national level structure to provide with policy and service delivery for the future and to educate the community to reduce violence against women and children. It included Australias National Research Organization for Womens Safety (ANROWS) , Our WATChandThe Line social marketing campaign.The second action plan included 5 important priorities such as driving whole community action to reduce violence, invigilating various types if cases of violence, introducing innovative systems and services to reduce the incidences, allow perpetrators interventions and continue collecting evidence bases in order for successful implementation of the policies ("Domestic and Family Violence factsheet | Department of Social Services, Australian Government", 2016). The policy given by Western Australian government are based on certain basic principles about violation of human right, community handling of issues, safety management and assurance, effective acknowledgement for proper maintenance of equity on gender, creed and caste etc. They also have unique policies for children to save them from the pressures of violence (Western Australian Government, 2016). The policy given by the Human services department looks over FDV helps the nation by active responsibly, providing support to the victims and their families, responding to them respectfully and including a collaborative approach for the proper guidance and maintenance of the cases and revival for the victims ("Family and domestic violence - Australian Government Department of Human Services", 2016). Explanation of effectiveness and evaluation of the policies: The policy produced by the department of Prime minister and cabinet have put forward principles that they have been found to have been fulfilled by them through a large number of different strategies which are not only well processed but also helps victims in every aspects not only mentally but also at a physical and financial level (Chapell Curtin, 2013). The large number of varieties of help lines provided had made the ways easier by which victims can seek for help (Department of Prime Minister and Cabinet, Australia, 2016). The various strategies of loans and various types of supports provided from the managerial levels such as allowances of various kinds has indeed effected in providing a workplace environment protective of the victims facing serious troubles in their personal troubles make the victims strong enough to face the adverse situations with courage and come out victorious. The plans given by the department of Social services has helped the nation to ensure the proper monitoring and reporting of cases of violence that has in turn been seen to reduce the prevalence of FDV ("Domestic and Family Violence factsheet | Department of Social Services, Australian Government", 2016). It has also committed to evaluate the planning of the policy on terms of the effectiveness and efficacy. Moreover, it had also ensured significant reduction of violence within the year 2022. The policy given by western Australia was evaluated with a n effective results in the fields of the performance indicator data such as the number of cases reported, police orders issued, perpetrators charges and others. Integrated interagency response has been found to be satisfactory on the associated outcomes. The Australian Bureau of Statistics Personal safety Survey also showed the number of reported cases to have reduced in Western Australia (Roberts, Chamberlain Delfabbro, 2015). Current policy Impact on equity, social justice and social determinants: Though the policy from prime ministerial department show equity in managing the victims on the basis if caste, creed religion and gender, however other policies although demand to be taking initiatives for the entire family members if any one faces violence, but specifically aims at protecting women and children. However, it is a fact that as the women and children are more vulnerable to abuses yet, policies for older citizens abuses were found no mention (Valentine Breckenridge, 2016). On the basis of caste and creed, equal laws and help lines are provide even to aboriginal and Non aborigines that show that policies are indeed effective as the provider of equal contribution to protect them from social injustice. Apart from the gender biasness with more preference in saving children and women policies were found to be more or less based on social justice and social determinants (Cross et al., 2012). Tensions and Alliances within the stakeholder groups: In order to discuss the various alliances, one must understand the importance of alliances among different stakeholder groups. For conducting the policies properly, a healthy alliance between the minister, policy makers, health professionals, counselors and the social workers are indeed necessary for the proper implementation of the policies (Fisher et al., 2016). However, unhealthy alliances between the perpetrators and the police authority or even members of the legislation and laws can result in bailing of the perpetrators that may increase the hardship on the victims and can be recognized as negative aspects of these alliances. Theses also give rise to several tensions among other stakeholders such as the victims and perpetrators. It often reflects in the victims minds that as the perpetrators are gaining bail every time, she also loses confidence on the laws and cases stop reporting abuses on her. Those, sometimes in many cases the victims do not only report violence against the m because of emotional attachment with the perpetrators and love for them that make them tolerate negligence and dominancy from the perpetrators. However, all these lead to unreported cases which in turn results in inefficiency on producing an effective plan for prevention of FDV and filing charges against perpetrators (Gilbert et al., 2012). Evidences of the benefits of policy Alternatives: In order to cut out the various tensions, a proper monitoring authority is very much important to be included in the policy so that any kind of corruptions and unethical behaviors can be totally excluded. This needs strong vigilance over each stakeholder involved so that the problems can be totally removed from grass root level. Care should be also made to make the victims confident to report every kind of abuses to the government so that the government can have the chances to alter the policies in favour of the victims (Fisher, 2013). Education should be made an important criteria in the country through various workshops and advertisements so that the women get knowledge about the policies and how to take the help of the policies to make their lives secure of themselves or of any victims around her. All these aspects should be included in a policy so that the recent concerns mentioned can also be included in the discussion arenas and can be included. An alternative strategy having p roper reporting of cases, good monitoring system, educating people of the ill effects and many others should also be included (Baird et al., 2014). Moreover another important criteria is the handling of each and every case with the perfect level of importance and sensitivity as per the demand of the cases because each and every case are different from each other and requires difference in urgency and different treatments. Therefore, case specific policies are important after partitioning the cases into different categories depending upon definite requirements and significance (Pain, 2014). Potential impact of the alternative policy on public health principles: The alternative policy that is designed including the loopholes present in the reviewed articles of the government would help to remove the tensions and the unethical and corrupted alliances between the stakeholders thereby providing policies that would ultimately help in establishing future that would be safe for every individuals irrespective of the caste , creed and the religions. The monitoring systems would thereby help to maintain a clean system that would reduce the levels FDV on the victims (Moser, 2012). Proper reporting of cases would help to produce statistical data that would help to analyze the policies both on the quantitative and qualitative results obtained from the policies and thereby course our future proposals and planning in the different regions of Australia. Conclusion: From analyzing the various policies put forward by the governments, one can easily analyses the features and positive aspects of the policies. However, it can well be understood that preparing a theoretical framework can act as a consort or a checklist which will help us to pinpoint the aspects that are missing and should be inculcated in the policies. In such cases, alternative policies are also found to be a good aid in solving the negative issues and prepare policies that would in turn create an environment in Australia safe from family and domestic violence. 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Retrieved 26 October 2016, from https://www.apsc.gov.au/__data/assets/pdf_file/0008/80954/domestic-family-violence-policy.pdf Domestic and Family Violence factsheet | Department of Social Services, Australian Government. (2016). Dss.gov.au. Retrieved 26 October 2016, from https://www.dss.gov.au/women/publications-articles/reducing-violence/domestic-and-family-violence Dunn, W. N. (2015).Public policy analysis. Routledge. Family and domestic violence - Australian Government Department of Human Services. (2016). Humanservices.gov.au. Retrieved 26 October 2016, from https://www.humanservices.gov.au/customer/subjects/family-and-domestic-violence Fisher, C. (2013). Changed and changing gender and family roles and domestic violence in African refugee background communities post-settlement in Perth, Australia.Violence against women,19(7), 833-847. Fisher, M., Milos, D., Baum, F., Friel, S. (2016). Social determinants in an Australian urban region: a complexitylens.Health promotion international,31(1), 163-174. Gilbert, R., Fluke, J., O'Donnell, M., Gonzalez-Izquierdo, A., Brownell, M., Gulliver, P., ... Sidebotham, P. (2012). Child maltreatment: variation in trends and policies in six developed countries.The Lancet,379(9817), 758-772. https://www.dcp.wa.gov.au/Documents/WA%20FDV%20Prevention%20Strategy%20to%202022.pdf Kettner, P. M., Moroney, R. M., Martin, L. L. (2016).Designing and managing programs: An effectiveness-based approach. Sage Publications. Larkin, H., Felitti, V. J., Anda, R. F. (2014). Social work and adverse childhood experiences research: Implications for practice and health policy.Social work in public health,29(1), 1-16. Lin, J. Y. (2012).New structural economics: A framework for rethinking development and policy. World Bank Publications. Moser, C. (2012).Gender planning and development: Theory, practice and training. Routledge. Pain, R. (2014). Everyday terrorism Connecting domestic violence and global terrorism.Progress in Human Geography,38(4), 531-550. Roberts, D., Chamberlain, P., Delfabbro, P. (2015). Women's experiences of the processes associated with the Family Court of Australia in the context of domestic violence: a thematic analysis.Psychiatry, Psychology and Law,22(4), 599-615. Sabatier, P. A., Weible, C. (Eds.). (2014).Theories of the policy process. Westview Press. Valentine, K., Breckenridge, J. (2016). Responses to family and domestic violence: supporting women?.Griffith Law Review,25(1), 30-44.